Penn State Science
Webb Miller

Webb Miller

Main Content

Professor of biology and computer science and engineering

506B Wartik
University Park, PA 16801
Phone: (814) 865-4551


  1. B.A., Mathematics, Whitman College, 1966
  2. M.S., Mathematics, University of Washington, 1967
  3. Ph.D., Mathematics, University of Washington, Seattle, 1969

Honors and Awards:

  1. Time Magazine 100 Most Influential People, 2009
  2. Senior Scientist Accomplishment Award, International Society of Computational Biology, 2009

Research Interests:

Algorithms and Software for Molecular Biology

Selected Publications:

Miller, W.,  et al. 2011. From the cover: Genetic diversity and population structure of the endangered marsupial Sarcophilus harrisii (Tasmanian devil). Proc. Natl. Acad. Sci. USA 108: 12348-12353.

Song, G., C. H. Hsu, C. Riemer, Y. Zhang, H. L. Kim, F. Hoffmann, L. Zhang, R. C. Hardison, E. D. Green, and W. Miller. 2011. Conversion events in gene clusters. BMC Evol. Biol. 11: 226.

Song, G., C. H. Hsu, C. Riemer, and W. Miller. 2011. Evaluation of methods for detecting conversion events in gene clusters. BMC Bioinformatics 12 (Suppl 1): 545.

Hsu, C.-H., Y. Zhang, R. C. Hardison, E. D. Green, and W. Miller. 2010. An effective method for detecting gene conversion events in whole genomes. J. Comput. Biol. 17: 1281-1297.

Song, G., Y. Zhang, T, Vinar, and W. Miller. 2010. CAGE: Combinatorial analysis of gene-cluster evolution. J. Comput. Biol. 17: 1227-1242.

Ratan, A., Y. Zhang, V. M. Hayes, S. C. Schuster, and W. Miller. 2010. Calling SNPs without a reference sequence. BMC Bioinformatics 11: 130.

Schuster, S. C., W. Miller, A. Ratan, et al. 2010. Complete Khoisan and Bantu genomes from southern Africa. Nature 463: 943-947.

Miller,W., et al. 2008 Sequencing the nuclear genome of the extinct woolly mammoth. Nature 456: 387-390.

Thomas, J. W., J. W. Touchman, [60 additional authors], W. Miller, and E. D. Green. 2003. Comparative analyses of multi-species sequences from targeted genomic regions. Nature 424: 788-793.

Janecka, J. E., W. Miller, T. H. Pringle, F. Wiens, A. Zitzmann, K. M. Helgen, M. S. Springer, and W. J. Murphy. 2007. Molecular and genomic data identify the closest living relative of primates. Science 318: 792-794.


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