Penn State Science
Webb Miller

Webb Miller

Main Content

Professor of Biology and Computer Science and Engineering

506B Wartik
University Park, PA 16802
Phone: (814) 865-4551


  1. B.A., Mathematics, Whitman College, 1966
  2. M.S., Mathematics, University of Washington, 1967
  3. Ph.D., Mathematics, University of Washington, Seattle, 1969

Honors and Awards

  1. Time Magazine 100 Most Influential People, 2009
  2. Senior Scientist Accomplishment Award, International Society of Computational Biology, 2009

Research Interests

Algorithms and Software for Molecular Biology

Selected Publications

Miller, W.,  et al. 2011. From the cover: Genetic diversity and population structure of the endangered marsupial Sarcophilus harrisii (Tasmanian devil). Proc. Natl. Acad. Sci. USA 108: 12348-12353.

Song, G., C. H. Hsu, C. Riemer, Y. Zhang, H. L. Kim, F. Hoffmann, L. Zhang, R. C. Hardison, E. D. Green, and W. Miller. 2011. Conversion events in gene clusters. BMC Evol. Biol. 11: 226.

Song, G., C. H. Hsu, C. Riemer, and W. Miller. 2011. Evaluation of methods for detecting conversion events in gene clusters. BMC Bioinformatics 12 (Suppl 1): 545.

Hsu, C.-H., Y. Zhang, R. C. Hardison, E. D. Green, and W. Miller. 2010. An effective method for detecting gene conversion events in whole genomes. J. Comput. Biol. 17: 1281-1297.

Song, G., Y. Zhang, T, Vinar, and W. Miller. 2010. CAGE: Combinatorial analysis of gene-cluster evolution. J. Comput. Biol. 17: 1227-1242.

Ratan, A., Y. Zhang, V. M. Hayes, S. C. Schuster, and W. Miller. 2010. Calling SNPs without a reference sequence. BMC Bioinformatics 11: 130.

Schuster, S. C., W. Miller, A. Ratan, et al. 2010. Complete Khoisan and Bantu genomes from southern Africa. Nature 463: 943-947.

Miller,W., et al. 2008 Sequencing the nuclear genome of the extinct woolly mammoth. Nature 456: 387-390.

Thomas, J. W., J. W. Touchman, [60 additional authors], W. Miller, and E. D. Green. 2003. Comparative analyses of multi-species sequences from targeted genomic regions. Nature 424: 788-793.

Janecka, J. E., W. Miller, T. H. Pringle, F. Wiens, A. Zitzmann, K. M. Helgen, M. S. Springer, and W. J. Murphy. 2007. Molecular and genomic data identify the closest living relative of primates. Science 318: 792-794.


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