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Timothy Reluga

Timothy Reluga

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Associate Professor of Mathematics and Biology

424 McAllister
University Park, PA 16802
Phone: (814) 865-3883


  1. Ph.D., Applied Mathematics, University of Washington, 2004
  2. B.S., Biology and Mathematics, Tufts University, 1998

Postdoc Training

  1. Postdoctoral Researcher, Los Alamos National Lab, 2006-2007

Research Interests

I'm an applied mathematician and member of the Center for Infectious Disease Dynamics (CIDD).  I am interested in the description, understanding, and prediction of the dynamics of populations, and the implications of biology for human communities. This currently includes work in ecology, epidemiology, immunology, evolution, medicine, human behavior, and economics.

Selected Publications

(see my NCBI bibliography also)


Unintended Consequences and the Paradox of Control: Management of Emerging Pathogens with Age-Specific Virulence, by Spencer Carran, Matthew Ferrari, and Timothy Reluga.
Accepted to PLoS Neglected Tropical Diseases , September, 2017.
Preprint PDF.


Provisioning of public health can be designed to anticipate public policy responses. By Jing Li, D. V. Lindberg, R. A. Smith, and T. Reluga.
Bulletin of Mathematical Biology, 79(1) 163-190, January, 2017.
DOI:10.1007/s11538-016-0231-8, Preprint PDF.


The importance of being atomic: ecological invasions as random walks instead of waves, by T. Reluga.
Theoretical Population Biology, December 2016, Volume 112, Pages 157-169.
DOI:10.1016/j.tpb.2016.06.002, Preprint PDF.


Resource distribution drives the adoption of migratory, partially migratory, or residential strategies. By T. Reluga and A. Shaw.
Theoretical Ecology, March, 2015.
DOI:10.1007/s12080-015-0263-y, Preprint PDF.


Population viscosity stops disease emergence by preserving local herd immunity. By T. Reluga and E. Shim.
Proceedings of the Royal Society B , 281(1796), 20141901, October 22, 2014.
DOI:10.1098/rspb.2014.1901, Preprint PDF.


Optimal migratory behavior in spatially-explicit seasonal environments. By Timothy Reluga and Allison Shaw.
Discrete and Continuous Dynamical Systems - Series B October 9, 2014, 19(10), 3359-3378. (Special issue in honor of Chris Cosner on the occasion of his 60th birthday, Submitted July, 2013).
DOI:10.3934/dcdsb.2014.19.3359, Preprint PDF


A reduction method for Boolean networks proven to conserve attractors. By Assieh Saadatpour, Reka Albert, and T. Reluga.
SIAM Journal on Applied Dynamical Systems, November, 2013, Volume 12, Issue 4, pp 1997-2011.
DOI:10.1137/13090537X, Preprint PDF.


Equilibria of an Epidemic Game with Piecewise Linear Social Distancing Cost By T. Reluga.
Bulletin of Mathematical Biology October 2013, Volume 75, Issue 10, pp 1961-1984.
DOI:10.1007/s11538-013-9879-5, Preprint PDF.


Games of age-dependent prevention of chronic infections by social distancing. By T. Reluga and J. Li.
Journal of Mathematical Biology, 2012.
DOI:10.1007/s00285-012-0543-8, Preprint PDF.


A general approach to population games with application to vaccination. By T. Reluga and A. Galvani.
Mathematical Biosciences, 230 (2): 67-78, April, 2011. (This paper received the 2013 Bellman Prize for best biannual paper)
DOI:10.1016/j.mbs.2011.01.003, Preprint PDF.